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1988,1986年的。1996年引述于 R. D. Guthrie. The Mammoth Steppe and the origin of Mongoloid and their dispersal. In Prehistoric Mongoloid Dispersal. Oxford Science Publications 1996。
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原文有 “as a general figure”的前提,当然非洲人肯定不都是这样的。
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Guthrie 引述的是:Harrison, G.A. et al., Human Biology, Princeton Univ. Press, 1988
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